Bombylius major (commonly named the large bee-fly, the dark-edged bee-fly or the greater bee fly) is a parasitic bee mimic fly. Bombylius major is the most common type of fly within the Bombylius genus. The fly derives its name from its close resemblance to bumblebees and are often mistaken for them.
Bombylius major exhibits a unique flight behavior known as “yawing” and plays a role in general pollination, without preference of flower types. The fly does not bite, sting, or spread disease. However, the fly uses this mimicry of bumblebees to its own advantage, allowing close access to host solitary bee and wasp nests to deposit its eggs. After hatching, the larvae find their way into the nests to parasitically feed on the grubs.
Flight
It has been discovered that the fly is capable of a unique behavior, which was discovered with the use of a high speed camera. In this behavior, the flies are seen to rotate around a vertical axis as they fly (this action is known as “yawing”). However, it is still unknown what can cause this behavior to be triggered and what purpose it serves, but a proposed explanation includes mating habits. Here is an illustration of “Yaw” in the context of an airplane…substitute the fly body with head facing forward (to left).
The Bombylius major bee-fly is a common, generalist floral pollinator, meaning that it does not give preference to one flower over another, instead pollinating a wide variety of plant families and species. The fly uses its proboscis to carry and transfer the pollen. The species is a dominant pollinator within its community, sometimes even pollinating up to two thirds of the local flowers. In addition, Bombylius major will visit and pollinate plants that attract few other species. Some types of flowers, for example Pulmonaria officinalis, will be almost exclusively pollinated by Bombylius major, with other species contributing a negligible amount to that plants pollination. Some flower species, such as Delphinium tricorne, are even specifically adapted to the fly in terms of color, shape, and form. If given the choice, Bombylius major will have a consistency in plant choice.
Flower Attraction
Long distance floral attraction is governed by optical sense, with color being the most important factor. The flies are typically more attracted to blue and violet colors, and occasionally yellow, over orange and pink. However, short distance floral attraction is based on the fly’s olfactory sense.
Sunbathing Activity
The fly is mostly active during day hours when the weather conditions are warm and sunny. Bombylius major is attracted to sunnier places and is more likely to pollinate these areas, with a larger average of flower visits in areas of higher amounts of sunshine. The fly will hide in the trees during the night and usually dart away from a cast shadow and occasionally hide in clean washing brought in fresh from the washing line and fly out causing unsettled behavior in the discoverer.
Flower Description
Claytonia caroliniana is a flowering, woodland perennial herb. It grows from March though June and is one of the earliest spring ephemerals. The plant grows from spherical underground tubers in light humus. They sprout and bloom before the tree canopy develops. Once the area is shaded, the plants whither leaving only the tuberous roots underground.
The plant is edible, but its usability is limited due to difficulty harvesting and the small quantities each plant produces. Its tuberous roots are edible and rich in starch and can be cooked or eaten raw. The leaves can be eaten as well. The tuberous roots are eaten by eastern chipmunks and white-footed mice.
Source: Wikipedia “Bombylius major ” and “Claytonia caroliniana. Direct quotations are in italics.
Copyright 2023 All Rights Reserved Michael Stephen Wills
Purple Trillium, a different species from the white, present different challenges. The purple blooms tend to dip down toward the ground. White flowers face upward toward the sky. My successful photographs of purple (Click me for another Purple Trillium posting) have the camera lower than the plant, say where there is a bank above the trail.
Shot from beneath, White Trillium project a hopeful air. Here is a comparison of the two species in the environmental and individual treatments.
Click photograph for larger image. To do this from WordPress Reader, you need to first click the title of this post to open a new page.
Pam and I ambled around the Arboretum for our Easter 2023 outing. We discovered this seeming lily growing from the muck along the Treman Woodland Walk. Scientific Name: Lysichiton camtschatcensis. Common names: Asian skunk cabbage, white skunk cabbage, Far Eastern swamp lantern or Japanese swamp lantern.
This is a plant found in swamps and wet woods, along streams and in other wet areas of the Kamchatka Peninsula, the Kuril Islands, Sakhalin and northern Japan. The common name “skunk cabbage” is used for the genus Lysichiton, which includes Lysichiton americanus, the western skunk cabbage, noted for its unpleasant smell. The Asian skunk cabbage is more variable: plants have been reported in different cases to smell disgusting, not at all, and sweet.
All photography using the IPhone 14 ProMax triple camera, raw format, edited on the phone.
In Japanese it is known as mizubashō (lit. “water-banana”) from a supposed similarity to the Japanese banana, a name with poetic rather than malodorous associations. It is not closely related to the true cabbage.
Copyright 2023 Michael Stephen Wills All Rights Reserved
Pam and I ambled around the Arboretum for our Easter 2023 outing. Found here growing outside native range, being the Appalachian Mountains from Georgia to southern Pennsylvania, the Table Mountain Pine is named after the landform, not a particular mountain.
All photography using the IPhone 14 ProMax triple camera, raw format, edited on the phone.
Its pinecones drew me to this scraggly, ungainly, poorly formed tree. All general mankind finds useful in the, scientific name, Pinus pungens, otherwise known as Hickory Pine, Prickly Pine and Mountain Pine, is to grind it up for pulp or chop it for tinder.
Last of the Mohicans
That said, the final scene of the 1992 film The Last of the Mohicans takes place in a nice Pinus pungens stand on a rocky mountaintop in North Carolina.
Personality
The tree has personality. Pinus pungens is the Lonesome Pine of the 1908 novel The Trail of the Lonesome Pine by John Fox, and popularized in the Laurel and Hardy film Way out West: “On the Blue Ridge Mountains of Virginia On the Trail of the Lonesome Pine” Several “Lonesome Pine” hiking trails have been waymarked in the Blue Ridge Mountains and elsewhere in the Appalachians.
Pinecone Bud
Pinus pungens prefers dry conditions and is mostly found on rocky slopes, favoring higher elevations, from 300–1,760 meters (980–5,770 ft) altitude. It commonly grows as single scattered trees or small groves, not in large forests like most other pines, and needs periodic disturbances for seedling establishment. The three tallest known ones are in Paris Mountain State Park, South Carolina; they are 26.85 to 29.96 meters (88 ft 1 in to 98 ft 4 in) tall.
Reference: “Pinus pungens” Wikipedia.
Copyright 2023 Michael Stephen Wills All Rights Reserved
Pam and I ambled around the Arboretum for our Easter 2023 outing. A type of sandstone popular with Cornell monument builders, called “Lenroc” after a mansion build by Cornell’s founder, was used for these benches built into the hillside of the FR Newman Arboretum. The views are more interesting than the bench, the arch of stone in midground in one photo.
All photography using the IPhone 14 ProMax triple camera, raw format, edited on the phone.
The stone is mined locally from surrounding hills. Calling it “Lenroc” (Cornell spelled backward) is a misnomer as the stone is mined widely throughout the region.
Feldspathic Greywacke
“Bluestone from Pennsylvania and New York is a sandstone defined as feldspathic greywacke. The sand-sized grains from which bluestone is constituted were deposited in the Catskill Delta during the Middle to Upper Devonian Period of the Paleozoic Era, approximately 370 to 345 million years ago…..
Glacial Landscape on an early spring day, Easter 2023
Textures
…The Catskill Delta was created from runoff from the Acadian Mountains (“Ancestral Appalachians”). This delta ran in a narrow band from southwest to northeast and today provides the bluestone quarried from the Catskill Mountains and Northeastern Pennsylvania. The term “bluestone” is derived from a deep-blue-colored sandstone first found in Ulster County, New York.”
You can feel the origin of this bluestone from these macros of two pavers from a monument bench.
Reference: “Bluestone” Wikipedia.
Copyright 2023 Michael Stephen Wills All Rights Reserved
These photographs are a continuation of blogs from two days’ exploration of Finger Rock Canyon of the Santa Catalina Mountains, southern Arizona. Here we explore the nature of the Saguaro blossom.
Saguaro flowers start as buds on the tip of the cactus body or arm. The specimen in the photograph below, growing in the yard of a foothills home on the border of federal land, is over 30 feet tall and, at the end of April 2011, buds are sprouting from every tip. Look closely for opening buds and full saguaro blossoms.
Flower buds grow only from some tips and around the center, along the sides, not from the point at the very end of the tip, from which the limb grows.
These buds first appeared mid-April and are here shown in the latter stages of maturity, prior to opening. Sometimes, the base of an arm weakens and the arm lowers close to the ground while remaining healthy. While descending the canyon I noticed this had happened to the arm of a particularly large specimen, an arm in full flower. This and the following photographs are from that arm.
I have read that each flower opens in the cool of the night and lasts only until the following afternoon. Here is a fully blossomed flower with a pair of opening buds.
And more, from a different view of the same arm.
A saguaro flower in full bloom, having opened the previous night. This flower will last a single day. It will wilt in the heat of a single afternoon and close. In this brief time, flying animals will pollinate it. You can see numerous honey bees on the flowers, in a previous blog, “Saguaro Flowers in Finger Rock Canyon.”
Our Winter Aconite started blooming around Valentines Day, February 14, 2023.
The following photograph is from the Apple IPhone 14 ProMax, raw format and perfected on the phone. The rest are from the Canon 5D Mark IV with the lens EF 100mm f/2.8 Macro USB.
As a spring ehemeral plant, its life cycle exploits the deciduous woodland canopy, flowering at the time of maximum sunlight reaching the forest floor, then completely dying back to its underground tuber after flowering.
All parts of the plant are poisonous when consumed by humans and other mammals because it contains cardiac glycosides.
The species name Eranthis hyemalis proclaims the early nature of its flowering both in the genus, “Eranthis” – “spring flower”, and species, “hyemalis” – winter flowering. The genus encompasses eight species, all early flowering winter aconite.
Reference: Wikipedia “Eranthis hyemalis” and “Eranthis.”
Copyright 2023 All Rights Reserved Michael Stephen Wills
I found crocuses blooming beneath our magnolia tree on our 14th anniversary, Monday, March 20, 2023.
The following photograph is from the Apple IPhone 14 ProMax, raw format and perfected on the phone. The rest are from the Canon 5D Mark IV with the lens EF 100mm f/2.8 Macro USB.
“Crocus is a genus of seasonal flowering plants in the family Iridaceae (iris family) comprising about 100 species of perennials growing from corms. They are low growing plants, whose flower stems remain underground, that bear relatively large white, yellow, orange or purple flowers and then become dormant after flowering. Many are cultivated for their flowers, appearing in autumn, winter, or spring. The flowers close at night and in overcast weather conditions.”
Here are partially closed blooms under the same magnolia tree on a very cold March 2020 afternoon.
“The crocus has been known throughout recorded history, mainly as the source of saffron. Saffron is obtained from the dried stigma of Crocus sativus, an autumn-blooming species. It is valued as a spice and dyestuff and is one of the most expensive spices in the world. Iran is the center of saffron production. Crocuses are native to woodland, scrub, and meadows from sea level to alpine tundra from the Mediterranean, through North Africa, central and southern Europe, the islands of the Aegean, the Middle East and across Central Asia to Xinjiang in western China. Crocuses may be propagated from seed or from daughter cormels formed on the corm, that eventually produce mature plants. They arrived in Europe from Turkey in the 16th century and became valued as an ornamental flowering plant.” Source: wikipedia “crocus”
Copyright 2023 All Rights Reserved Michael Stephen Wills
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